Monday, March 4, 2019
Effects of Migration and Other Evolutionary Processes on Allele
Effects of migration and sepa straddle evolutionary processes on allelomorph frequence and fitness Life originated from a common ancestor and due to various mechanisms of evolution, the genotype of organisms has changed. vicissitude, migration, inherited impetus and plectrum ar natural processes of evolution that affect agenttic diversity. Mutations atomic number 18 spontaneous changes in genomic sequences (Robert, et al. , 2006) it is one of the processes that influence allele frequency. A pas seul elicit either entertain a positive, negative or a achromatic effect on an organisms fitness.When organisms of the same species shew different phenotypes, the organism is polymorphic for that particular trait. A beneficial sportsman that gives evolve to polymorphic traits wad cleanse the chance of survival. For example, the grove snail, Cepaea nemoralis, is not equal to(p) for the rich polymorphism of its shell. A sportsman in the locus trusty for colour produces differe nt shell colours, ranging from discolor, pink, white and brownish (Ozgo, 2005). Snails with brown shells atomic number 18 found in beechwoods where the soil is dark.Snails with brown shells be able to mask with the soil, thus avoiding being detected by predators (Jones, et al, 1977). As a resolving of avoiding predation, the frequency of alleles that code for brown shells result increase. However, according to the hitchhiking model, fixation of a beneficial mutation go forth decrease the diversity at cereb target loci (Chevin, et al. , 2008). If a new mutation increases the fitness of members of a particular species, a strong selective sweep on allele frequency exit result to very few haplotypes existing in the race.The frequency of alleles that are positively selected and those that are closely linked will increase, but the former(a) alleles will decrease. A mutation can be neutral, having neither a beneficial effect nor a negative effect. However, several(prenominal) mutations are deadly because they overhear a negative effect on fitness. The accumulation of pernicious mutations and the prevention of recombination flash back the fitness of individuals (Mullers ratchet). Experiment carried out on a inner and sexual yeast strains showed that sexually reproducing parts of the genome improved survival than asexually reproducing parts (Zeyl and Bell, 1997).Asexual strains decreased over beat because of Mullers ratchet. On the contrary, sexual strains were able to stop the build-up of deleterious mutation due to recombination between chromosomes. Mutation in collagen-I gene is another example of lethal mutation trim back fitness. Collagen is a group of naturally occurring proteins found in animals, it is one of the major(ip) components of blood vessels. An experiment carried out on mouse embryonic fore cells showed that mutation in collagen-I gene impairs the function of collagen-I (Lohler, et al. 1984). During the experiment, 13 embryos died be cause a mutation in mouse collagen-I gene caused the major blood vessels to rupture. According to terra firma extract model, because a deleterious mutation reduces the fitness of individuals, deleterious mutations are selected against (Innan and Stephan, 2003) this will decrease the allele frequency of a macrocosm. Genetic rate is a stochastic process that refers to the fluctuations of genotype frequencies (Maynard, 1998) alleles are either strict or permanently lost from the population.Due to the randomness of the process, genetic drift can eliminate beneficial alleles that could birth improved survival. Genetic drift can also eliminate lethal alleles from a population and therefore improve survival rate. Genetic drift has larger effect on subtle populations than a large population (Maynard, 1998) this is because the rate of allele fixation or elimination is faster in a small population compared to a large population. Moreover, population bottleneck is an evolutionary proces s that increases the effect of genetic drift it involves random events that prevent species from reproducing (van-Heerwaarden, et al. 2008). Population bottleneck decreases allele frequency and it reduces a populations capability to adapt to new environmental pressures. For example, the current cheetah populations have low genetic diversity caused by a demographic bottleneck that occurred 10,000 years ago (Charruau, et al. , 2011). The surviving cheetah populations are not representative of the original cheetah population because they have less(prenominal) variation (founder effect). Due to low genetic diversity and less variant skills, the modern cheetah population is close to extinction. Natural selection is another evolutionary process that changes allele frequency.Organisms with advantageous alleles survive and reproduce, change magnitude the frequency of the advantageous alleles. Individuals with disadvantageous alleles do not survive or reproduce and therefore the frequenc y of the disadvantageous alleles is reduced or eliminated from the population (William and Michael, 2003). Biston betularia (peppered moths) is a common example used to demonstrated natural selection (Saccheri, et al. , 2008). Before the industrial revolution, non-melanic peppered moths avoided predators by camouflaging with lichen-covered trees.Their ability to camouflage improved the rate of survival which increased the frequency of non-melanic alleles. Melanic peppered moths were not able to camouflage with the lichen trees, as a result, melanic moths were detected and predated by the song thrushes. This decreased the frequency of alleles that gave acquire to melanic peppered moths. However, during the industrial revolution period, symbiotic lichens living on trees were killed because smog and vulgarism were released when coal and other materials were burnt.As a consequence of the tree trunks decent more visible, non-melanic peppered moths were more susceptible to predation be cause they were unable to camouflage with the trees. The ability to camouflage helped melanic moths to survive and reproduce, changing the population allele frequency from broadly non-melanic alleles to mostly melanic alleles (Saccheri, et al. , 2008). Migration of species from one place to another can increase the rate of gene flow. Gene flow is the transfer of gene from one population to another (William and Michael, 2003) it changes the allele frequency of a population.The effect of migration on the gene pool of a population depends on the rate of migration. Various studies have shown that migration rate is not the same for all species (Tajima, 1990). Species with low migration rate will have less desoxyribonucleic acid polymorphism and species with high migration rate will have more polymorphic alleles (Tajima, 1990). The usefulness of plant migration, which increases the chance of crisscross between plant species, can be demonstrated by examining the adaptation skills of Iri s species. Iris nelsonii is a species of hybrid origin, with traces of I. fulva, I. hexagona and I. revicaulis. I. nelsonii picked up characteristics that are not present in the parent population. For example, I. nelsnii can grow in homophile(a) wet conditions whereas the parents can either grow in sunny juiceless conditions or wet and shady conditions (Taylor, et al, 2011). Given that I. nelsonii can survive in challenging environments, the allele frequency of the advantageous traits will increase. Furthermore, another benefit of gene flow through means of hybridization can be demonstrated by analyzing the genetic variation of Tragopogan species. Hybridization between T. enigmatic and T. pratensis produces T. iscellus, an allotetraploid that has multiple enzymes needed for various biochemical path ways (Tate, et al. , 2006). Hybridisation enabled T. miscellus and T. pratensis to survive because they were able to exploit the gene pool of both parents. However, migration can also have negative effects on survival. Given that I. nelsonii will exist in niches that parents cannot live in, gene flow between the hybrid and its progenitors will be reduced. If I. nelsonii does not have alleles that can resists infection caused by parasites, an outbreak of a pathogenic disease can wipe out the entire I. nelsonii species.Although some evolutionary processes eliminate alleles from a population, multiple alleles can be well-kept through frequency-dependent balancing selection (Matessi and Schneider, 2009). In negative frequency-dependent selection, the fitness of a phenotype increases as it becomes less common. An example of negative frequency-dependent selection is in the elusion of Cepaea nemoralis. C. nemoralis are regularly predated by song thrush birds called Turdus philomelos. These birds have a search pattern whereby it persists in targeting the most abundant morph, even if other morphs are available (Bond, 2007).If snails with yellow shells are common, then th ese snails will be eaten by song thrushes. As a result, the frequency of alleles that code for yellow shells will decrease. The fitness of other morphs such as pink, white and brown shells will increase because song thrushes would not search for rare dark morphs. In conclusion, the four fundamental processes of evolution, mutation, genetic drift, natural selection and migration (gene flow), alters allele frequencies in populations. The consequences on survival fluctuate. Occasionally, altering allele frequency gives rise to traits that increases fitness.However, changing allele frequencies can also give rise to phenotypes that reduce fitness. Word count 1390 Grade A- My essay is easy to read and follow. I have given evidences and interpreted them where possible. I also gave examples from animals and plants to show that I have done outside reading. All of the points that were made are applicable as they ultimately answer4 the question e. g. whether the evolutionary processes increa se of decrease allele frequency and fitness References Bond, AB, 2007. The evolution of color polymorphism crypticity searching images, and apostatic selection.Annual round Of Ecology Evolution And Systemic, 38, pp. 489-514. Charruau, P. , Fernandes, C. , Orozco-ter Wengel, P. , Peters, J. , Hunter, L. , Ziaie, H. , Jourabchian, A. , Jowkar, H. , Schaller, G. , Ostrowski, S. , Vercammen, P. , Grange, T. , Schlotterer, C. , Kotze, A. , Geigl, EM. , Walzer, C. and Burger, PA. (2011). Phylogeography, genetic structure and population divergence time of cheetahs in Africa and Asia evidence for long-term geographic isolates. Molecular Ecology, 20, pp. 706-724. Chevin, LM. , Billiard, S. and Hospital, F. (2008).Hitchhiking both ways Effect of two interfering selective sweeps on linked neutral variation. Genetics, 180, pp. 301-316. Innan, H. and Stephan, W. (2003). Distinguishing the Hitchhiking and Background Selection Models. Genetics, 165, pp. 2307-2312. Jones, J. S. , Leith, B. N. and Rawlings, P. (1977). Polymorphism in cepaea a problem with in any case many solutions. Annual Reviews In Ecology And Systematics, 8, pp. 10914. Lohler, J. , Timpl, R. and Jaenisch, R. (1984). Embronic lethal mutation in mouse collagen-I gene causes rupture of blood-vessels and is associated with erythropoietic and mesenchymal cell-death.Cell, 38, pp. 597-607. Matessi, C. and Schneider, KA. (2009). Optimization under frequency-dependent selection. divinatory Population Biology, 76, pp. 1-12. Maynard, S. J. (1998). Evolutionary genetics. 2nd edition. new York Oxford University Press. Ozgo, M. (2005). Cepaea nemoralis (L. ) in southeastern Poland joining of morph frequencies with habitat. Journal Of Molluscan Studies, 71, pp. 93-103. Saccheri, IJ. , Rousset, F. , Watts, PC. , Brakefield, PM. and Cook, LM. (2008). Selection and gene flow on a diminishing cline of melanic peppered moths.Proceedings Of The National Academy Of sciences Of The United States Of America, 105, pp. 16212-16 217. Tajima, F. (1990). Relationship between migration and DNA polymorphism in a local-population. Genetics, 126, pp. 231-234. Tate, JA. , Ni, ZF. , Scheen, AC. , Koh, J. , Gilbert, CA. , Lefkowitz, D. , Chen, ZJ. , Soltis, PS. , Soltis, DE. (2006). Evolution and expression of homeologous loci in genus Tragopogon miscellus (Asteraceae), a recent and reciprocally formed allopolyploid. Genetics, 173, pp. 1599-1611. Taylor, SJ. , Willard, RW. , Shaw, JP. , Dobson, MC. and Martin, NH. (2011).Differential response of the homoploid hybrid species iris diaphragm nelsonii (iridaceae) and its progenitors to abiotic habitat conditions. American Journal Of Botany, 98, pp. 1309-1316. van Heerwaarden, B. , Willi, Y. , Kristensen, TN. and Hoffmann, AA. (2008). Population bottlenecks increase analogue genetic variance but do not break a selection limit in rain forest Drosophila. Genetics, 179, pp. 2135-2146. William, S. K. and Michael, R. C. (2003). Concepts of genetics. 7th edition. New Jersey P earson Education Limited. Zeyl, C. and Bell, G. (1997). The advantage of sex in evolving yeast populations. Nature, 388, pp. 465-468.
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